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            <gco:CharacterString>What: soil carbon; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: biomass burning; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea water temperature; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea water salinity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: northward ocean heat transport; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: dissolved inorganic carbon; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: Delta 14C; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: phosphate; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: diazotroph; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: dissolved organic nitrogen; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: diazotroph; Material: sea water</gco:CharacterString>
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            <gco:CharacterString>What: diazotroph; Material: sea water</gco:CharacterString>
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            <gco:CharacterString>What: snow depth; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: upward sea water velocity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: upward sea water velocity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: grazing; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: nitrogen fixation; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: remineralization; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: surface air temperature; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: carbon dioxide; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea ice cover; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: insolation; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: vegetation carbon; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: soil carbon; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: vegetation carbon; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: canopy height; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: dissolved inorganic carbon; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: zooplankton; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: sea water y velocity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: specific humidity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: evaporation; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: top-of-atmosphere outgoing longwave radiation; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: alkalinity; Material: sea surface</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: litter carbon; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: vegetation carbon; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: sea water salinity; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: 14C; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: organic matter; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: dissolved organic carbon; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: streamflow; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: virtual salt flux; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: surface pressure; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: soil moisture; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: ice temperature; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: surface downward x stress; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: surface downward y stress; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea water potential temperature; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea water x velocity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: snow volume; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: surface net downward shortwave radiation; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: top-of-atmosphere net downward shortwave radiation; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: dissolved inorganic carbon; Material: sea surface</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: primary productivity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea water temperature; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: nitrate; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: phytoplankton; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: phytoplankton; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: organic matter; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: meridional overturning streamfunction; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: runoff; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea ice y velocity; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: carbon dioxide; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: delta 13C; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: Delta 14C; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: 14C; Material: sea surface</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: primary productivity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: primary productivity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: primary productivity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: leaf area index; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: dissolved inorganic carbon; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: 13C; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: oxygen; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: organic matter; Material: sea water</gco:CharacterString>
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            <gco:CharacterString>What: dissolved organic nitrogen; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: zooplankton; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: zooplankton; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: aragonite saturation state; Material: sea surface</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: wind speed; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: ice sheet area; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: calcite; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: sea water x velocity; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: grazing; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: mortality; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: phytoplankton; Material: sea water</gco:CharacterString>
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            <gco:CharacterString>What: meridional overturning streamfunction; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: barotropic streamfunction; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: dissolved organic nitrogen; Material: sea surface</gco:CharacterString>
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            <gco:CharacterString>What: dissolved organic nitrogen; Material: sea surface</gco:CharacterString>
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            <gco:CharacterString>What: soil respiration; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: 14C; Material: sea water</gco:CharacterString>
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            <gco:CharacterString>What: snowfall; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: northward wind speed; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea ice thickness; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: grazing; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: mortality; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea ice volume; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: surface net upward longwave radiation; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: top-of-atmosphere net downward radiation; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea surface temperature; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea surface salinity; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: dissolved organic carbon; Material: sea surface</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: biomass burning; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: soil carbon; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: soil respiration; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: biomass burning; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: temperature tendency; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: oxygen; Material: sea water</gco:CharacterString>
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            <gco:CharacterString>What: pH; Material: sea surface</gco:CharacterString>
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            <gco:CharacterString>What: calcite saturation state; Material: sea surface</gco:CharacterString>
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            <gco:CharacterString>What: sedimentation rate; Material: null</gco:CharacterString>
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            <gmd:MD_KeywordTypeCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#MD_KeywordTypeCode" codeListValue="theme">theme</gmd:MD_KeywordTypeCode>
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                            <gmd:URL>https://www.ncei.noaa.gov/products/paleoclimatology/paleoenvironmental-standard-terms-thesaurus</gmd:URL>
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            <gmd:MD_KeywordTypeCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#MD_KeywordTypeCode" codeListValue="dataCentre">dataCentre</gmd:MD_KeywordTypeCode>
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ABSTRACT SUPPLIED BY ORIGINATOR: Changes in the ocean iron cycle during the Last Glacial Maximum (LGM) have been proposed to help explain the low atmospheric CO2 from this period. Previous global modeling studies, however, have only considered changes in aeolian iron fluxes, although it is known that sedimentary and hydrothermal fluxes are important iron sources for today's ocean. Here we evaluate the effect of preindustrial-to-LGM changes in atmospheric dust, sedimentary, and hydrothermal fluxes on the ocean's iron and carbon cycles in a global coupled biogeochemical circulation model. Considering variable iron solubility we find that higher dust fluxes into the ocean in the LGM increase productivity and enhance the biological pump, sequestering carbon in deep waters and lowering CO2 by 4 ppm. The effect is countered by a decrease in sedimentary release of iron due to lower sea level, which increases CO2 by 15 ppm. In an upper-limit estimation of Southern Ocean surface iron fertilization, assuming a 10 times higher solubility for this region, combined with changes in sedimentary release we find an increase in deep dissolved inorganic carbon concentrations, that could potentially reduce atmospheric CO2 by 13 ppm. A five-fold increase in hydrothermal sources of iron during the LGM may have decreased atmospheric CO2 by an additional 6 ppm. We conclude that when addressing LGM-preindustrial changes in the ocean iron cycle, not only surface but also sedimentary and hydrothermal fluxes have to be taken into account.</gco:CharacterString>
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