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                <gco:CharacterString>Parameter or Variable: BIOMASS (calculated);
Units: tons/square kilometer;
Observation Category: model output;
Sampling Instrument: Not applicable;
Sampling and Analyzing Method: Ecopath with Ecosim (EwE) version 6.4.4 was used to conduct sensitivity and hindcast simulations. Fish species were aggregated into functional groups and all biomass data was converted to metric tons per square kilometer and used as input data for the EwE models for Laysan and French Frigate Shoals.

For the sensitivity assessment, four perturbation scenarios were simulated for a time span of 40 years (2010 through 2049): (a) baseline, (b) altered predator abundance, (c) altered monk seal prey abundance, and (d) altered primary productivity. The sensitivity assessment datasets include forecast simulations with monthly output for the last 5 years of the 40-year simulation run (2044-2049). Monthly output includes mean biomass of a variety of marine organisms, such as seabirds, sharks, fish, turtles, plankton, and algae.

For the hindcast simulations, historical stressors to monk seal population dynamics were evaluated by selecting the best-fit model for monk seal biomass time series (1998-2015). The hindcast simulation datasets for 1998 through 2015 include simulated monk seal and bottomfish biomass factoring a few key external drivers (fishing with time series based on commercial fishery data and additional mortality simulated with a constant removal of monk seals of varied intensities being the most important).

Time series observation data of monk seal biomass and benthic bottomfish biomass, as well as catch time series of bank sharks (only for FFS), benthic and demersal bottomfish, bank jacks, and macroheterotrophs (lobsters) were loaded into Ecosim for model fitting. Best-fit of model-simulated output data and observational data was based on the least square fitting criterion and summed the squared deviations. Additionally, the Akaike's information criterion (AIC) was calculated taking into account the small sample size.

Refer to Weijerman et al (2017) for complete details. Weijerman M, Robinson S, Parrish F, Polovina J, Littnan C (2017) Comparative application of trophic ecosystem models to evaluate drivers of endangered Hawaiian monk seal populations. Mar Ecol Prog Ser 582:215-229. https://doi.org/10.3354/meps12320;
Data Quality Method: Model dynamics were validated with a hindcast simulation (1998-2015). Time series data of monk seal biomass and bottomfish biomass were loaded into Ecosim for validation and to fine-tune the model by adjusting vulnerability values and so minimize the residuals between predicted and observed time series of monk seal and bottomfish biomass using a least-square fitting criterion. We manually adjusted vulnerability values of interest based on the results from the perturbation scenarios and restricted them to ecological meaningful values (i.e., preferably between 1 and 3 with 10 as maximum value).</gco:CharacterString>
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