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            <gco:CharacterString>What: northward ocean heat transport; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: dissolved inorganic carbon; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: Delta 14C; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: phosphate; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: diazotroph; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: dissolved organic nitrogen; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: diazotroph; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: diazotroph; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: snow depth; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: upward sea water velocity; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: upward sea water velocity; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: grazing; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: nitrogen fixation; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: remineralization; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: evaporation; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: dissolved organic phosphorus; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: specific humidity; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: top-of-atmosphere outgoing longwave radiation; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: vegetation carbon; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: soil carbon; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: vegetation carbon; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: canopy height; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: dissolved inorganic carbon; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: zooplankton; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: sea water y velocity; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: iron; Material: sea surface</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: snow volume; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: surface net downward shortwave radiation; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: top-of-atmosphere net downward shortwave radiation; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: alkalinity; Material: sea surface</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: litter carbon; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: vegetation carbon; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: salinity; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: 14C; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: organic matter; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: dissolved organic carbon; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: streamflow; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: virtual salt flux; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: surface pressure; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: soil moisture; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: ice temperature; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: surface downward x stress; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: surface downward y stress; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: sea water potential temperature; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: sea water x velocity; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: iron; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: carbon dioxide; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: delta 13C; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: dissolved inorganic carbon; Material: sea surface</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: primary productivity; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: sea water temperature; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: salinity; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: nitrate; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: phytoplankton; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: phytoplankton; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: organic matter; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: meridional overturning streamfunction; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: runoff; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: sea ice y velocity; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: dissolved organic phosphorus; Material: sea surface</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: Delta 14C; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: 14C; Material: sea surface</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: primary productivity; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: primary productivity; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: primary productivity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: leaf area index; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: dissolved inorganic carbon; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: 13C; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: oxygen; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: organic matter; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: dissolved organic nitrogen; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: zooplankton; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: zooplankton; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: aragonite saturation state; Material: sea surface</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: wind speed; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: ice sheet area; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: calcite; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea water x velocity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea water y velocity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: grazing; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: mortality; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: grazing; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: export; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: phytoplankton; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: precipitation; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: sensible heat flux; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: planetary albedo; Material: null</gco:CharacterString>
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          <gmd:keyword>
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            <gco:CharacterString>What: soil respiration; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: nitrate; Material: sea water</gco:CharacterString>
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            <gco:CharacterString>What: meridional overturning streamfunction; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: carbonate ion; Material: sea surface</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: latitude; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: sea ice cover; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea ice x velocity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: barotropic streamfunction; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: excretion; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: mortality; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: denitrification; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: export; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea ice volume; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: surface net upward longwave radiation; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: top-of-atmosphere net downward radiation; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea surface temperature; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea surface salinity; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: oxygen; Material: sea surface</gco:CharacterString>
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            <gco:CharacterString>What: dissolved organic nitrogen; Material: sea surface</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: dissolved organic nitrogen; Material: sea surface</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: soil respiration; Material: null</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: 14C; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: alkalinity; Material: sea water</gco:CharacterString>
          </gmd:keyword>
          <gmd:keyword>
            <gco:CharacterString>What: relative sea level; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: snowfall; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: northward wind speed; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: sea ice thickness; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: grazing; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: mortality; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: iron; Material: sea water</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: dissolved organic carbon; Material: sea surface</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: biomass burning; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: soil carbon; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: soil respiration; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: biomass burning; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: temperature tendency; Material: null</gco:CharacterString>
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            <gco:CharacterString>What: northward ocean salt transport; Material: null</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: oxygen; Material: sea water</gco:CharacterString>
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            <gco:CharacterString>What: pH; Material: sea surface</gco:CharacterString>
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            <gco:CharacterString>What: calcite saturation state; Material: sea surface</gco:CharacterString>
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          <gmd:keyword>
            <gco:CharacterString>What: carbon dioxide; Material: sea surface</gco:CharacterString>
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            <gco:CharacterString>What: eastward wind speed; Material: null</gco:CharacterString>
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                        <gmd:CI_OnlineResource>
                          <gmd:linkage>
                            <gmd:URL>https://www.ncei.noaa.gov/products/paleoclimatology/paleoenvironmental-standard-terms-thesaurus</gmd:URL>
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            <gmd:MD_KeywordTypeCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#MD_KeywordTypeCode" codeListValue="dataCentre">dataCentre</gmd:MD_KeywordTypeCode>
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ABSTRACT SUPPLIED BY ORIGINATOR: The role of glacial ocean circulation and iron fertilization in the sequestration of atmospheric CO2 are not well understood. Here we use a physical-biogeochemical model constrained by carbon and nitrogen isotopes to explore the effects of Atlantic Meridional Overturning Circulation (AMOC) and Southern Ocean iron fertilization on the distributions of these isotopes and ocean carbon storage. We find that a surprisingly weak (6-9 Sv) and shallow AMOC maximizes carbon storage and best reproduces the glacial sediment data. Increasing the atmospheric soluble iron flux in the Southern Ocean intensifies export production, carbon storage, and improves agreement with reconstructions. Our best fitting model suggests that a weak, shallow AMOC and enhanced iron fertilization conspired to enhance carbon storage in the glacial ocean.</gco:CharacterString>
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